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Tusk was an original recruit of the Riders of the Storm, who served Apocalypse both when he attempted to reclaim his Ship from X-Factor, and later during their fight with the Inhumans on the Moon.
Afterwards, the Dark Riders carried on Apocalypse's Darwinian dogma on their own, fighting, testing, and often killing mutants who didn't meet their standards. Cyclops was their only known target who proved to be one of the strong.
Tusk was initially seen with the Dark Riders when they were adopted by Genesis, the self-appointed heir to Stryfe and Apocalypse; however, he disappeared early into Genesis' leadership and so wasn't present when Wolverine mauled and eviscerated the Dark Riders that remained.
After mutants began falling ill and dying due to the effects of the Terrigen Mist spread out into the atmosphere, the Dark Riders returned. They began murdering mutant healers, believing that their powers were a violation of the natural order, their reasoning being that mutants had had their chance at survival, and that delaying their extinction was counter to the natural order.
They attracted the attention of Magneto and his X-Men, who managed to secure Triage before the Riders could get to him. While the X-Men attacked the Riders' lair in the hope of taking them out preemptively, the Riders set a trap for the X-Men and left to attack Genosha, where Magneto was protecting Triage alone.
They managed to critically injure Triage, despite heavy interference from Magneto, but when the rest of the X-Men arrived to back him up, Magneto was able to overpower the Riders. He left them dead or critically injured on the island, which he then detonated and sunk into the Indian Ocean with a series of explosive charges, finishing off Tusk and the rest of the Riders.
Enhanced Form: Tusk has the power to grow a monstrous, massive exo-form that possesses superhuman strength and durability, as well as additional chitinous armor material and two bull-like horns on his back.
- Mini-Clone Production: Tusk can also grow smaller versions of himself that sprout from his back and operate independently while maintaining a psychic link with the main body. His clones also possess enhanced strength and durability, as well as enhanced agility.
- Tusk's smaller selves are occasionally referred to as Underlings (capital "U"). This is a running error dating back to the X-Cutioner's Song when Stryfe monologued about dispatching Tusk's "underlings" in combat, but in context it was not meant to be an official "codename" for the creatures.
- 18 Appearances of Tusk (Earth-616)
- Media Tusk (Earth-616) was Mentioned in
- 9 Images featuring Tusk (Earth-616)
- Quotations by or about Tusk (Earth-616)
- Character Gallery: Tusk (Earth-616)
- The Unofficial Handbook to the Marvel Universe
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- ↑ 1.0 1.1 All-New Official Handbook of the Marvel Universe #3
- ↑ 2.0 2.1 X-Man #46
- ↑ X-Factor #65-66
- ↑ X-Factor #68
- ↑ X-Men Vol 2 #14-16
- ↑ X-Force #17
- ↑ X-Factor #86
- ↑ X-Men Vol 2 #21
- ↑ X-Men Vol 2 #23
- ↑ Cable #17-19
- ↑ Wolverine Vol 2 #100
- ↑ Uncanny X-Men Vol 4 #1
- ↑ Uncanny X-Men Vol 4 #4
- ↑ Uncanny X-Men Vol 4 #5
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