Joanne Tumolo was a member of the Cat people, living disguised as a human scientist doing research on motor response at the University of Chicago. Malcolm Donalbain helped her finance a private experiment she was conducting, to allow any woman to totally fulfill her physical and mental potential, eventually turning her lab assistant Greer Grant into the superhero the Cat.

When Grant was shot with an alpha radiation weapon while defending Dr. Tumulo from a kidnapping attempt by Hydra agents, the doctor arranged for the Cat People to transform Greer into one of them. She became Tigra[1]. After her transformation, Tumolo brought Tigra to Professor Leon to expose her to a process that would allow her to take human form.[2]

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